Univoltine, with an extended flight period. Females from early April to mid October; males from early July to mid October.
Label data from a few Spanish specimens show that females are active from (at least) March to May (Ortiz-Sanchez & Pauly, 2017)
Parasites and predators R.C.L. Perkins (1923) recorded Sphecodes hyalinatus and S. ferruginatus as cleptoparasites.
On the continent, Sphecodes crassus is cited as a cleptoparasite (Söderman & Leinonen, 2003)