Distribution; world
Central and south eastern Europe, eastwards into Asia Minor and the Caucasus:
AT, BG, CZ, GR, GR-AN, GR-DD, GR-KR, HR, HU, IT, IT-SI, MD, MK, PL, RO, RS, RU, RU-EU, SK, TR, UA, UA-KY, YU, AM, IR
Conservation status; world
Conservation status: CZ - RDB0 (RE), EU - RDB-DD (Nieto et al, 2014)