Distribution; world
Regarded as a Pontic species, Ranging from western Europe eastwards across Asia Minor to the Caucasus:
AT, BG, CZ, DE, ES, FR, GR, GR-AN, GR-DD, HR, HU, IT, LT, PL, RO, RU, SL, SK, TR, UA, YU, GA
Conservation status; world
Conservation status: CZ - RDB2 (EN), D1E - RDB2 (EN), DE2 - RDB1 (EN), EU - RDB-DD (Nieto et al, 2014)