Excavator: Ground. K.M. Guichard (1970) observed females entering nesting burrows on a firm, sparsely vegetated, sandy slope. On the Continent, the species is considered to nest solitarily (Dylewska, 1987). S.P.M. Roberts and G.R. Else noted a large dispersed nesting aggregation in a sloping exposed bank in calcareous grassland on Salisbury Plain (GB)