Oligolectic on Fabaceae (e.g. Onobrychis, Medicago, Melilotus, Lotus, Vicia), an important pollen source are flowers of the Hedysareae (e.g. Onobrychis). Several pollen samples from the same locality were found to contain small amounts of Boraginaceae pollen, which was probably not actively collected, indicating that flowers of the Boraginaceae are sometimes used as nectar sources (Özbek, 1979b; Özbek and Zanden, 1992a; A. Müller, unpublished, based on 52 pollen samples from 11 different localities).