Lowland/upland vegetation zones including sclerophylous biotopes (0-350m a.s.l.)(GR)(Standfuss et al., 2011)
This species was found on roadsides, fallow lands, and shores of rivers (Nipru river, Ukraine). It prefers xeric habitats from open steppes and coastal areas of Black Sea and Azov Sea to mountainous region of Caucasus (Talysh Mt.) and semi-arid region of central Anatolia (Konya basin)(EU)