Calcareous grassland, heathland edge, open woodland (GB)
Radish (Raphanus sativus ‘Omega’) fields (BG)
Humid habitats (woods edge, valleys, nutrient-rich meadows and grasslands, ruderal field borders and fences, and plantations) up to 1,800 m in the Swiss Alps (Osytshnjuk 1977, Dylewska 1987, Calabuig and Madsen 2009)(EU)