A species of Beech (Fagus sylvatica) forests (SL) (Gogala, 1999)
Woodland edges, sparse grassy areas and waste place (CH)
Extensive upland pasture with Scabiosa columbaria (Dipsacaceae) (FR) (Pauly & Belval; 2017)
Beech forests, in sub-Mediterranean and pre-alpine regions in addition to woodland edges, sparse grassy areas and waste places (EU)