Tall sward calcareous grasslands and vegetated dune systems supporting strong populations of the forage plants (GB)
Lowland/upland vegetation zones including sclerophylous biotopes (0-350m a.s.l.)(GR)(Standfuss et al., 2011)(GR)
Dry grasslands with plentiful Knautia arvensis (NL)
Valley slopes with thermophilous vegetation (BG)
Moist meadows and pastures next to forest habitats, for example in Romania and Ukraine. It is also found in open grasslands, calcareous and sandy soils in inland and coastal habitats, such as fixed dunes in the United Kingdom (Else 2001). They also appear on flower-rich road sides and alpine grasslands up to 2,000-2,500 m asl. (EU)