Dry meadows and other dry grasslands, river levees and man-made habitats (DE)
Floristically rich ares in phryganic habitats (GR-AN)
Lowland/upland vegetation zones including sclerophylous biotopes (0-350m a.s.l.)(GR)(Standfuss et al., 2011)
Moist habitats such as forests and grassland edges, alpine meadows and grasslands in the valleys (Osytshnjuk 1977), and it has also been found on dry meadow slopes. In the mountains it rises up to 2,000 m asl (Pereval Bichenekskiy, Caucasus, Azerbaijan; unpublished data Osytshnjuk, V. Radchenko pers. comm. 2014)(EU)