Serpentine rocky steppic grassland (CZ)
Mainly in the mountains on grasslands and shrublands at altitudes up to 2,300 m asl in the Transcaucasus (Osytshnjuk unpublished data V. Radchenko pers. comm. 2014), 1,900 m asl in Greece (Gusenleitner 1984) and 1,800 m asl in Macedonia (Warncke 1973a). In Moravia (Czech Republic) it is a steppic species (Přidal and Vesely 2011).(EU)