Rough coastal grassland and cliffs, heathland, calcareous grassland, country lanes and moorland edge (GB)
Lowland/upland vegetation zones including sclerophylous biotopes (0-350m a.s.l.)(GR)(Standfuss et al., 2011)
Dry and humid habitats (grassland slopes and meadows) up to 1,440 m asl in Turkey (Warncke 1969a, Osytshnjuk 1977)(EU)