Steppic grasslands in the Crimea (UA)
Bound mainly to forest, forest edge or forest clearings (IR, RS, RO, GR) (Baker, 1996, Knechtel, 1955, Reinig, 1972).
Oak forest edge (SK)(Sima & Smetana, 2009)
Mixed Carpinus/Populus/Fagus forest edge (RS)(Roberts, pers. obs.)